Ao

A beetle who was once infected and now lives freed from the Radiance. He is busy trying to put his life back together after living as a zombie for an undisclosed period of time. When Hallownest still stood, he was a commander of great skill who used his one wing as a built-in blade. He is still incredibly skilled, but he rarely gets to use his blade anymore.

He is a gynandromorph, meaning that his body is literally split between male and female. As a result, his horns are un-even and he only has one wing.